Muhammad Akhtar, Pakistani biochemist (Batala, Punjab 23 February 1933 –

Defined the complete substrate stereochemistry of at least a dozen enzymes

Elucidated the mode and site of binding of the retinal chromophore in bovine rhodopsin (1969-83)

First to describe the structure of bovine rhodopsin in terms of trans-membrane segments (1978)

With J.N. Wright discovered that P-450 group of enzymes catalyze a diverse range of generic reactions at a single active site (1982)